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I've tracked this down to a bad subsetting of the fac matrix, which results in an empty toDrop object.
However, there's another deeper problem. Even after making sure that the formula terms where speciesCode and cover appear are correctly identified, the way the function is written results in completely dropping (logAge + cover + speciesCode | ecoregionGroup), instead of keeping (logAge | ecoregionGroup)...
reprex
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